- Dave as Manny
- Simon as Sid
- Alvin as Diego
- Theodore as Scrat
- Eleanor as Scratte
- Clarie as Ellie
- Rodney & Fender (from Robots) as Crash & Eddie
- Marge (from The Simpsons) as Peaches
- Homer (from The Simpsons) as Ethan
- Brittany as Shira
- Jeanette as Granny
- Ian as Captain Gutt
- Zoe as Raz
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