- Yar (from Disney's Dinosaur) as The Lorax
- Django (from Ratatouille) as Once-ler
- Fillmore as Young Once-ler
- Lightning McQueen as Ted
- Sally as Audrey
- Mater as Pipsqueak
- Emily (from TTTE) as Granny Norma
- Lizzie as Ted's Mom
- Professor Z (Cars 2) as O'Hare
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