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- Sonic the Hedgehog as Ariel
- Amy Rose as Prince Eric
- Sally Acorn as Sebastian
- Cosmo the Plant as Scuttle
- Cream the Rabbit as Flounder
- Rouge the Bat as King Triton
- Dr Eggman as Ursula
- Miles "Tails" Prower as Melody
- Wave as Hammerhead shark
- Knuckles the Echidna, Espio the Chalmeleon, Shadow the Hedgehog, and Silver the Hedgehog as Triton's Daughter by Ariel's sister
- Vector the Crocodile as Carlotta
- Vanilla the Rabbit as Adult Flounder
- Sonia as Grimsby