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- Emily as Rayman (Emily and Rayman are both the main stars and the main heroes)
- Thomas as Ly the Fairy
- Edward as Uglette (Edward and Uglette are both husbands to Molly and Globox)
- Diesel 10 as Mr. Dark
- Molly as Globox (Molly and Globox are both wifes to Edward and Uglette and good friends with Emily and Rayman)
- Daisy as Admiral Razorbeard
- Flora as Murfy
- Madge as Clark
- Tillie (from The Little Engine That Could) as Sam the Snake
- Mavis as Polokus
- Misty, May, Dawn, and Iris (from Pokemon) as the Teensies
- Toad as Carmen
- and more