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- Manny as Alex
- Diego as Marty
- Macrauchenia as Melman
- Granny as Gloria
- Sid as King Julien
- Aardvark as Maurice
- Scrat as Mort (both really small)
- Elmo (from Sesame Street) and Fievel (from An American Tail) as Mason and Phil
- Big Bird (from Sesame Street) as Kowalski
- Leo (from Little Einsteins) as Skipper
- Danny (from CDD) as Rico
- Boo Boo Bear (from Yogi Bear) as Private
- Scratte as Marlene