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  • Donald as The Lorax
  • Arthur as The Once-ler
  • Thomas as Young Once-ler
  • James as Ted
  • Madge as Ted's Mom
  • Isobella as Audrey
  • Whiff as Pipsqueak
  • Daisy as Grammy Norma
  • Bulgy as Mr. O'Hare
  • Douglas as Fat Bar-ba-loot
  • Splatter and Dodge as The once-ler's Brothers

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