- Snowball as Alvin
- Luke as Simon
- Junior as Theodore
- Junior and Luke's Owner as Brittany
- Chloe as Jeanette
- Tessa Yeager (from Transformers: Age of Extintion) as Eleanor
- Mayor Lionheart as Dave
- Grammy Norma (from Dr. Seuss' The Lorax) as Ashley
- Kevin as Benson
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