- Timon as Alvin (both are leaders but are vain)
- Adult Simba as Simon (both intelligent)
- Pumbaa as Theodore
- Mufasa as Dave
- Scar as Ian
- Sarabi as Claire
- Marlene (from The Penguins of Madagascar) as Brittany
- Adult Nala as Jeanette
- Ma as Eleanor
- Zira as Zoe
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