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- Rodney Copperbottom - Thomas/Edward/Sunshine (from TUGS)/Himself/Gordon/Diesel/Donald/Douglas/Bertie/Butch/Arthur/Stanley/Stepney/Percy/Toby/Henry/Duck/Toad/Hector/Horrid Lorries/Bill/Ben/D261/Spencer/Murdoch/City of Truro/The Fat Controller/Cranky/Bert (RWS)/James
- Herb Copperbottom - Odie (from Garfield)/Ten Cents (from TUGS)/Big Mac (from TUGS)/Salty/Boco/Diesel/Diesel 10/D261/James/Edward/Gordon/Sonic (from Sonic)/Uncle Chuck (from Sonic)/Antoine (from Sonic)/Homer (from The Simpsons)/Donald/Kevin/Victor/Luke/Duke/Whiff/Douglas/Derek/Dennis/Thomas/Sir Topham Hatt/Himself
- Lydia Copperbottom - Pearl (from TUGS)/Sally Seaplane (From TUGS)/Lillie Lightship (from TUGS)/Marge (from The Simpsons)/Henry/Gordon/James/Thomas/Percy/Mavis/Daisy/Emily/Duchess of Hamilton/Sally (from Sonic)/Amy (from Sonic)/Queen Aleena (from Sonic)/Breezie (from Sonic)/Bunnie (from Sonic)/Lady Hatt/Rosie/Herself
- Cappy - Lady/Emily/Pearl (from TUGS)/Herself/Madge/Rosie/Annie/Clarabel/Thomas/Mavis/Molly/Caroline/Elizabeth/Judge Judy
- Phineas T. Ratchet - Spencer/Diesel/D261/Bluenose (from TUGS)/Diesel 10/Murdoch/Thomas/Gordon/James/Percy/Toby/Donald/Douglas/Bill/Ben/Mavis/Salty/Duke/Freddie/Arthur/Oliver/Stepney/Duck/Himself/D199/Old Stuck Up/Sir Topham Hatt
- Mr. Gunk - Warrior (from TUGS)/Mr. Krabs (from SpongeBob)/Knuckles (from Sonic)/Thomas/James/Donald/Douglas/Percy/Duck/Bear/Mr. Percival/Santa/The Fire Truck/Toby/Oliver/Stepney/Toby/Trucks/Daisy/Chinese Dragon/Himself/Diesel/Diesel 10/George/Hector/Homer (from The Simpsons)
- Diesel - Edward/Diesel/Diesel 10/D261/Bulgy/Thomas/James/Oliver/Duck/Stepney/Percy/Edward/Donald/Douglas/Ada/Jane/Mabel/Mr. Conductor/Sir Topham Hatt's Car/Murdoch
- Garbage Robots - Coaches/Trucks/Blue Trucks/Skarloey/Peter Sam/Duncan/Season 1 Trucks/Weasels (from Who Framed Roger Rabbit)/Themselfs/Thomas Villains/Horrid Lorries/NG Coaches/The TUGS Characters/Disney Villains/Scratch, Grounder, Coconuts and Badniks (from Sonic)/Z-Stacks (from TUGS)