- Timothy as Alvin Seville
- Fritz as Simon Seville
- Charles as Theodore Seville
- Yoko as Brittany Miller
- Lily as Jeanette Miller
- Nora as Eleanor Miller
- Mr. Raccoon as Dave Seville
- Mrs. Jenkins as Miss Miller
- Mrs. Raccoon as Vinny/Claire Wilson
- Henry as Uncle Harry/Ian Hawke
- Miss Appleberry as Miss Stone/Dr. Rubin
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